Hosted by one of the oldest and prominent universities in Europe, Jagiellonian University, the meeting will welcome hundreds of researchers, educators, and students from around the world, all of whom are immersed in some aspect of symbiosis. Held every three years and organized by the International Symbiosis Society, the Congress is one of the most unique gatherings of life science research specialists in the world. As symbiotic systems encompass and even dominate many phyla and most domains and kingdoms, it is a venue wherein an expert in coral-dinoflagellates will exchange ideas, results, methods, and perspectives with a mycorrhizae or lichen specialist. Those in the vast field of insect symbioses interact with those in the legume-nitrogen fixing realm. In this sense, the term “symbiosis,” applies very well to the Congress experience, as extended exchanges, long-term relationships, and new lineages of thought emerge from this diverse human community.